Deflected Alexis Sanchez free-kick gave Arsenal lead at Everton but Seamus Coleman equalised on the cusp of half-time with a header. Ashley Williams headed Everton in front with four minutes to play.
Premier League champions will rue their squandered chances as Marc Pugh fired Bournemouth winner against Leicester City
Everton 2-1 Arsenal
Arsenal missed the chance to go top of the Premier League as Everton came from behind to secure a first victory since October.
The Gunners, who had won their previous three games, were rewarded for an impressive start as Alexis Sanchez's free-kick deflected in off Ashley Williams.
Everton equalised just before half-time when Seamus Coleman glanced home Leighton Baines' excellent cross.
The Toffees, encouraged by a vocal crowd, dominated thereafter and Williams powered home a late header to complete the turnaround.
Home captain Phil Jagielka was sent off in the dying minutes for a second bookable offence and Arsenal had two shots cleared off the line in a frantic finale.
But they could not prevent a first league defeat since their opening match of the season, leaving them three points behind leaders Chelsea, who have a game in hand.
Everton, meanwhile, are up to seventh, four points behind fifth-placed Tottenham.
How many times have you seen Arsene Wenger this way? He Sat in between his assistants, wincing his displeasure, unable to understand why things aren't going to plan.
Arsene Wenger is an open book when Arsenal squander a chance to make progress and here he was again, scowling and complaining, lamenting the loss of precious points.
How crucial will this be in the final reckoning? It is a question the Frenchman will ask himself over and over in the next few days.
Bournemouth 1-0 Leicester City
Another German goalkeeper in a flap at Bournemouth and another night when it seemed entirely possible that the most romantic story in all sports will end with a devastating anti-climax.
The more games there are like this, and the longer this hangover goes on, the more likely it seems that Leicester City will wake up rubbing their head in the Championship and wondering where they left their trousers.
A side that cantered through a season of tremendous achievement are now so frail that they followed a win against Manchester City by wilting here, offering precious little in attack and almost as little at the back.
Bournemouth moved to the highest league position in their history as Leicester's miserable away run from home continued with defeat at Vitality Stadium.
The champions paid for a lacklustre first half, where they created clear openings for Jamie Vardy and Robert Huth, but surrendered the ball all too easily.
Eddie Howe's side saw 70% of the ball before the break and Marc Pugh scored for the hosts on his first league start of the season with a controlled drive from 18 yards out.
Leicester improved but ran into a stubborn home defence and goalkeeper Artur Boruc produced a key save to deny Leonardo Ulloa from six yards in the final minute.
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